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Heyer, W. R. 2014. Morphological analyses of frogs of the Leptodactylus latrans complex (Amphibia, Leptodactylidae) from selected localities in South America. Proceedings of the Biological Society of Washington, 126(4): 369-378. doi:10.2988/0006-324X-126.4.369

Sá, Rafael O. d., Grant, Taran, Camargo, Arley, Heyer, W. R., Ponssa, Maria L. and Stanley, Edward 2014. Systematics of the Neotropical Genus Leptodactylus Fitzinger, 1826 (Anura: Leptodactylidae): Phylogeny, the Relevance of Non-molecular Evidence, and Species Accounts. South American Journal of Herpetology, 9: S1-S100. doi:10.2994/SAJH-D-13-00022.1

Heyer, W. R. 2014. Morphological analyses of frogs of the Leptodactylus latrans complex (Amphibia, Leptodactylidae) from selected localities in South America. Proceedings of the Biological Society of Washington, 126(4): 369-378. doi:10.2988/0006-324X-126.4.369

Heyer, W. R. and Heyer, Miriam M. 2013. Systematics, distribution, and bibliography of the frog Leptodactylus insularum Barbour, 1906 (Amphibia: Leptodactylidae). Proceedings of the Biological Society of Washington, 126(3): 204-233. doi:10.2988/0006-324X-126.3.204

Heyer, W. R. and Heyer, Miriam M. 2012. Systematics, distribution, and bibliography of the frog Leptodactylus validus Garman, 1888 (Amphibia: Leptodactylidae). Proceedings of the Biological Society of Washington, 125(3): 276-294. doi:10.2988/12-04.1

Heyer, Miriam M., Heyer, W. R. and de Sa, Rafael O. 2012. Leptodactylus pentadactylus. Catalogue of American Amphibians and Reptiles, 887: 1-48.

Heyer, W. R. and de Sá, Rafael O. 2011. Variation, Systematics, and Relationships of the Leptodactylus bolivianus Complex (Amphibia: Anura: Leptodactylidae). Smithsonian Contributions to Zoology, 635: 1-58. doi:10.5479/si.00810282.635.1

Furness, Andrew I., McDiarmid, Roy W., Heyer, W. Ronald and Zug, George R. 2010. Oviduct modifications in foam-nesting frogs, with emphasis on the genus Leptodactylus (Amphibia, Leptodactylidae). South American Journal of Herpetology, 5(1): 13-29. doi:10.2994/057.005.0102

Lavilla, Esteben O., Langone, Jose A., Caramaschi, Ulisses, Heyer, W. Ronald and De Sá, Rafael Omar 2010. The identification of Rana ocellata Linnaeus, 1758. Nomenclatural impact on the species currently known as Leptodactylus ocellatus (Leptodactylidae) and Osteopilus brunneus (Gosse, 1851)(Hylidae). Zootaxa, 2346: 1-16.

Laura Ponssa, Maria, Heyer, W. Ronald and De Sá, Rafael Omar 2010. Phylogeny of the genus Leptodactylus (Anura, Leptodactylidae) based on characters of morphology and mode of life. Cladistics, 26(2): 222

Heyer, W. R., Heyer, Miriam M. and De Sá, Rafael Omar 2010. Leptodactylus syphax. Catalogue of American Amphibians and Reptiles, 868: 1-9.

Heyer, W. R., Heyer, Miriam M. and De Sá, Rafael Omar 2010. Leptodactylus savagei. Catalogue of American Amphibians and Reptiles, 867: 1-19.

Fernández, Miguel, Cole, Daniel, Heyer, W. Ronald, Reichle, Steffen and De Sá, Rafael Omar 2009. Predicting Leptodactylus (Amphibia, Anura, Leptodactylidae) distributions: Broad-ranging versus patchily distributed species using a presence-only environmental niche modeling technique. South American Journal of Herpetology, 4(2): 103-116. doi:10.2994/057.004.0202

Heyer, Miriam H., Heyer, W. Ronald and De Sá, Rafael Omar 2009. Bibliography of the frogs of the Leptodactylus clade - Adenomera, Hydrolaetare, Leptodactylus, Lithodytes (Amphibia, Anura, Leptodactylidae). Volume 2. Indices. Smithsonian Herpetological Information Service, 2: 1-167. doi:10.5479/si.23317515.2.1

Heyer, Miriam H., Heyer, W. Ronald and De Sá, Rafael Omar 2009. Bibliography of the frogs of the Leptodactylus clade - Adenomera, Hydrolaetare, Leptodactylus, Lithodytes (Ampbibia, Anura, Leptodactylidae). Volume 1. References. Smithsonian Herpetological Information Service, 1: 1-677. doi:10.5479/si.23317515.1.1

Heyer, W. R. and Barrio-Amorós, César Luis 2009. The advertisement calls of two sympatric frogs, Leptodactylus lithonaetes (Amphibia: Anura: Leptodactylidae) and Pristimantis vilarsi (Amphibia: Anura: Strabomantidae). Proceedings of the Biological Society of Washington, 122(3): 282-291. doi:10.2988/09-02.1

Heyer, W. R. and Giaretta, Ariovaldo A. 2009. Advertisement calls, notes on natural history, and distribution of Leptodactylus chaquensis (Amphibia: Anura: Leptodactylidae) in Brasil. Proceedings of the Biological Society of Washington, 122(3): 292-305. doi:10.2988/08-42.1

Camargo, Arley, Heyer, W. Ronald and De Sá, Rafael Omar 2009. Phylogeography of the frog Leptodactylus validus (Amphibia: Anura): Patterns and timing of colonization events in the Lesser Antilles. Molecular Phylogenetics and Evolution, 53: 571-579.

Heyer, W. R. 2008. Can long-established scientific societies survive the digital age?. Journal of the Washington Academy of Sciences, 94(2): 17-32.

Heyer, W. R., Heyer, M. M. and De Sá, Rafael Omar 2008. Leptodactylus cunicularius. Catalogue of American Amphibians and Reptiles, 845: 1-5.

Bolaños, F., Castro, F., De la Riva, I., Grant, T., Hedges, B., Heyer, W. Ronald, Ibáñez D., Roberto, La Marca, E., Lavilla, E., Leite Silvano, D., Lötters, S., Parra Olea, G., Reichle, S., Reynolds, Robert P., Rodríguez, L., Santos Barrera, G., Scott, N., Ubeda, C., Veloso, A., Wilkinson, M. and Young, B. 2008. Amphibians of the Neotropical Realm. In: Stuart, S. N., Hoffmann, M., Chanson, J. S., Cox, N. A., Berridge, R. J., Ramani, P. and Young, B. E., Threatened Amphibians of the World. Barcelona: Lynx Edicions, pp.92-105.

Ponssa, M. L. and Heyer, W. Ronald 2007. Osteological characterization of four putative species of the genus Adenomera (Anura: Leptodactylidae), with comments on intra- and interspecific variation. Zootaxa, 1403: 37-54.

Camargo, Arley, De Sá, Rafael Omar and Heyer, W. Ronald 2006. Phylogenetic analyses of mtDNA sequences reveal three cryptic lineages in the widespread neotropical frog Leptodactylus fuscus (Schneider, 1799) (Anura, Leptodactylidae). Biological Journal of the Linnean Society, 87(2): 325-341. doi:10.1111/j.1095-8312.2006.00581.x

Heyer, W. R. and Heyer, Miriam Muedeking 2006. Leptodactylus knudseni . Catalogue of American Amphibians and Reptiles, 807: 1-12.

Heyer, W. R. and Heyer, Miriam Muedeking 2006. Leptodactylus natalensis . Catalogue of American Amphibians and Reptiles, 808: 1-5.

Heyer, W. R. and Heyer, M. M. 2006. Leptodactylus knudseni. Catalogue of American Amphibians and Reptiles, 807: 1-12.

Heyer, W. R. and Heyer, M. M. 2006. Leptodactylus natalensis. Catalogue of American Amphibians and Reptiles, 808: 1-5.

Heyer, Miriam M. and Heyer, W. Ronald 2006. Leptodactylus fragilis (Brocchi) white-lipped thin-toed frog. Catalogue of American Amphibians and Reptiles, 830: 1-26.

Heyer, W. R. and Scott, Norman J., Jr. 2006. The advertisement call of Leptodactylus laticeps (Amphibia, Anura, Leptodactylidae): predatory aural luring?. Herpetological Natural History, 9(2): 189-194.

Heyer, W. R., Caramaschi, Ulisses and De Sá, Rafael Omar 2006. Case 3323. Rana ocellata Linnaeus, 1758 (currently Leptodactylus ocellatus; Amphibia, Anura): proposed conservation of usage of the specific name by the designation of a neotype. Bulletin of Zoological Nomenclature, 63(3): 184-186.

Heyer, W. R. 2006. Obituary: in celebration of A. Stanley Rand (1932 - 2005). Phyllomedusa, 5(1): 8-10.

Yanek, Keneth, Heyer, W. Ronald and De Sá, Rafael Omar 2006. Genetic resolution of the enigmatic lesser Antillean distribution of the frog Leptodactylus validus (Anura, Leptodactylidae). South American Journal of Herpetology, 1(3): 192-201. doi:10.2994/1808-9798(2006)1[192:GROTEL]2.0.CO;2

Heyer, W. R., De Sá, Rafael Omar and Muller, Sarah 2005. On the enigmatic distribution of the Honduran endemic Leptodactylus silvanimbus (Amphibia: Anura: Leptodactylidae). In: Donnelly, Maureen A., Crother, Brian I., Guyer, Craig, Wake, Marvalee H. and White, Mary E., Ecology and evolution in the tropics: a herpetological perspective. Chicago: University of Chicago Press, pp.81-101.

Heyer, W. R. 2005. Leptodactylus fragilis Brocchi, 1877. White-lipped frog. In: Lannoo, M., Amphibian Declines. The Conservation Status of United States Species. University of California Press, pp.500-501.

Hayek, Lee-Ann C. and Heyer, W. Ronald 2005. Determining sexual dimorphism in frog measurement data: integration of statistical significance, measurement error, effect size and biological significance. Anais Da Academia Brasileira De Ciencias, 77(1): 45-76.

Heyer, W. R. and Murphy, James B. 2005. Declining Amphibian Populations Task Force. In: Lannoo, Michael, Amphibian declines: the conservation status of United States species. Berkeley, CA: University of California Press, pp.17-21.

Heyer, W. R., De Sá, Rafael Omar and Rettig, A. 2005. Sibling species, advertisement calls, and reproductive isolation in frogs of the Leptodactylus pentadactylus species cluster (Amphibia, Leptodactylidae). Herpetologica Petropolitana.Russian Journal of Herpetology, 12: 35-39.

Caramaschi, Ulisses, De Sá, Rafael Omar and Heyer, W. Ronald 2005. Common names for the frog genus Leptodactylus (Amphibia, Anura, Leptodactylidae). Herpetological Review, 36(2): 119-120.

De Sá, Rafael O., Heyer, W. Ronald and Camargo, Arley 2005. A phylogenetic analysis of Vanzolinius Heyer, 1974 (Amphibia, Anura, Leptodactylidae): taxonomic and life history implications. Arquivos do Museu Nacional, 63(4): 707-726.

Heyer, W. R. and Crombie, Ronald I. 2005. Leptodactylus lauramirimae, a distinctive new species of frog (Amphibia: Anura: Leptodactylidae) from RondA'nia, Brazil. Proceedings of the Biological Society of Washington, 118(3): 590-595.

De Sá, Rafael O., Heyer, W. Ronald and Camargo, A. 2005. Are Leptodactylus didymus and L. mystaceus phylogenetically sibling species (Amphibia, Anura, Leptodactylidae)?. Herpetologica Petropolitana.Russian Journal of Herpetology, 12: 90-92.

Brandão, Reuber A. and Heyer, W. Ronald 2005. The complex calls of Leptodactylus pustulatus (Amphibia, Anura, Leptodactylidae). Amphibia-Reptilia, 26(4): 566-570.

Heyer, W. R. 2005. Variation and taxonomic clarification of the large species of the Leptodactylus pentadactylus species group (Amphibia: Leptodactylidae) from Middle America, northern South America, and Amazonia. Arquivos de Zoologia Sao Paulo, 37(3): 269-348.

Heyer, W. R. and Heyer, Miriam Muedeking 2004. Leptodactylus furnarius Sazima and Bokermann cerrado oven frog. Catalogue of American Amphibians and Reptiles, 785: 1-5.

Heyer, W. R. 2004. Paulo Emilio Vanzolini. Copeia, 2004(1): 184-189.

Heyer, W. R. and Reid, Yana R. 2003. Does advertisement call variation coincide with genetic variation in the genetically diverse frog taxon currently known as Leptodactylus fuscus (Amphibia: Leptodactylidae)?. Anais da Academia Brasileira de Ciencias, 75(1): 39-54.

Heyer, W. R. 2003. Ultraviolet-B and Amphibia. (Viewpoint). Bioscience, 53(6): 540-541.

Heyer, M. M., Heyer, W. Ronald, Spear, S. and De Sá, Rafael Omar 2003. Leptodactylus Mystacinus (Burmeister) Mustached Frog. In: Catalogue of American Reptiles and Amphibians. Society for the Study of Amphibians and Reptiles, pp.767.1-767.11.

Heyer, W. R., Huelsbergen, K. J., Wittmann, Ch, Papaja, S. and Christen, O. 2003. Field related organisms as possible indicators for evaluation of land use intensity. Agriculture Ecosystems and Environment, 98(1-3): 453-461.

Heyer, M. M., Heyer, W. Ronald, Spear, S. and De Sá, Rafael Omar 2003. ’Leptodactylus mystacinus.’. Catalogue of American Amphibians and Reptiles, 767: 1-11.

Heyer, W. R. and Junca, Flora A. 2003. Leptodactylus caatingae, a new species of frog from eastern Brazil (Amphibia: Anura: Leptodactylidae). Proceedings of the Biological Society of Washington, 116(2): 317-329.

Heyer, W. R. and Caramaschi, Ulisses 2002. Comment on the proposed conservation of the specific name of Leptodactylus chaquensis Cei, 1950 (Amphibia, Anura). Bulletin of Zoological Nomenclature, 59(1): 44-45.

Heyer, W. R. and Heyer, Miriam Muedeking 2002. Leptodactylus elenae Heyer. Catalogue of American Amphibians and Reptiles, 742: 742

Heyer, W. R., De Sá, Rafael Omar and Muller, Sarah 2002. Leptodactylus silvanimbus McCranie, Wilson, and Porras. Catalogue of American Amphibians and Reptiles, 743: 743

Heyer, W. R. 2002. Leptodactylus fragilis, the valid name for the Middle American and northern South American white-lipped frog (Amphibia: Leptodactylidae). Proceedings of the Biological Society of Washington, 115(2): 321-322.

Wynn, Addison H. and Heyer, W. Ronald 2001. Do Geographically Widespread Species of Tropical Amphibians Exist? An Estimate of Genetic Relatedness Within the Neotropical Frog Leptodactylus fuscus (Schneider 1799) (Anura Leptodactylidae). Tropical Zoology, 14: 255-285.

Hayek, Lee-Ann C., Heyer, W. Ronald and Gascon, Claude 2001. Frog morphometrics: A cautionary tale. Alytes Paris, 18(3-4): 153-177.

Carey, Cynthia, Heyer, W. Ronald, Wilkinson, John, Alford, Ross A., Arntzen, J. W., Halliday, Tim, Hungerford, Laura, Lips, Karen R., Middleton, Elizabeth M., Orchard, Stan A. and Rand, Austin Stanley 2001. Amphibian declines and environmental change: use of remote-sensing data to identify environmental correlates. Conservation Biology, 15(4): 903-913.

Heyer, W. R. and Heyer, Miriam Muedeking 2001. Leptodactylus lithonaetes Heyer. Catalogue of American Amphibians and Reptiles, 723: 723

Heyer, W. R. and Morato de Carvalho, Celso 2000. The enigmatic advertisement call of Eleutherodactylus ramagii (Amphibia: Anura: Leptodactylidae). Amphibia-Reptilia, 21(1): 117-121.

Heyer, W. R. 2000. The lectotype of Cystignathus hylodes Reinhardt and Lutken, 1862. Journal of Herpetology, 34(1): 150-153.

Heyer, W. R. and Morato de Carvalho, Celso 2000. Calls and calling behaviour of the frog Leptodactylus natalensis (Amphibia: Anura: Leptodactylidae). Proceedings of the Biological Society of Washington, 113(1): 284-290.

Heyer, W. R. and Thompson, A. S. 2000. Leptodacylus rugosus Noble. In: Catalogue of American Amphibians and Reptiles. Society for the Study of Amphibians and Reptiles, pp.708.1-708.5.

Heyer, W. R., Coddington, Jonathan A., Kress, W. John, Acevedo-Rodríguez, Pedro, Cole, Daniel, Erwin, Terry L., Meggers, Betty J., Pogue, Michael G., Thorington, Richard W., Jr., Vari, Richard P., Weitzman, Marilyn J. and Weitzman, Stanley H. 1999. Amazonian biotic data and conservation decisions. Ciência e Cultura Sao Paulo, 51(5-6): 372-385.

Heyer, W. R. and Munoz, Anna M. 1999. Validation of Eleutherodactylus crepitans Bokermann, 1965, notes on the types and type locality of Telatrema heterodactylum Miranda-Ribeiro, 1937, and description of a new species of Eleutherodactylus from Mato Grosso, Brazil (Amphibia: Anura: Leptodactylidae). Proceedings of the Biological Society of Washington, 112(1): 1-18.

Heyer, W. R. 1999. A new genus and species of frog from Bahia, Brazil (Amphibia: Anura: Leptodactylidae) with comments on the zoogeography of the Brazilian campos rupestres. Proceedings of the Biological Society of Washington, 112(1): 19-39.

Heyer, W. R. 1998. The relationships of Leptodactylus diedrus (Anura, Leptodactylidae). Alytes Paris, 16(1-2): 1-24.

Kress, W. J., Heyer, W. Ronald, Acevedo-Rodríguez, Pedro, Coddington, Jonathan A., Cole, D., Erwin, Terry L., Meggers, Betty J., Pogue, Michael G., Thorington, Richard W., Jr., Vari, Richard P., Weitzman, Marilyn J. and Weitzman, Stanley H. 1998. Amazonian biodiversity: Assessing conservation priorities with taxonomic data. Biodiversity and Conservation, 7(12): 1577-1587.

Savage, Jay M. and Heyer, W. Ronald 1997. Digital webbing formulae for anurans: a refinement. Herpetological Review, 28(3): 131

Heyer, W. R. 1997. Geographic variation in the frog genus Vanzolinius (Anura: Leptodactylidae). Proceedings of the Biological Society of Washington, 110(3): 338-365.

Halliday, Timothy R. and Heyer, W. Ronald 1997. The case of the vanishing frogs. Technology review, 100(4): 56-63.

Heyer, W. R., Garcia Lopez, Jose M. and Cardoso, Adao J. 1996. Advertisement call variation in the Leptodactylus mystaceus species complex (Amphibia: Leptodactylidae) with a description of a new sibling species. Amphibia-Reptilia, 17(1): 7-31.

Heyer, W. R., De Sá, Rafael Omar, McCranie, James R. and Wilson, Larry David 1996. Leptodactylus silvanimbus (Amphibia: Anura: Leptodactylidae): natural history notes, advertisement call, and relationships. Herpetological Natural History, 4(2): 169-174.

Heyer, W. R. and Morales, Victor R. 1995. The advertisement call of the leptodactylid frog Leptodactylus griseigularis. Amphibia-Reptilia, 16(1): 91-92.

Heyer, W. R. 1995. South American rocky habitat Leptodactylus (Amphibia: Anura: Leptodactylidae) with description of two new species. Proceedings of the Biological Society of Washington, 108(4): 695-716.

Heyer, W. R. 1995. Methodische Untersuchungen zur Erfassung von Schadinsekten in Bohnenbestanden (Phaseolus vulgaris L.). Archives of Phytopathology and Plant Protection, 29(6): 533-553.

Heyer, W. R. and Gascon, Claude 1995. Collection notes and call characteristics for Ischnocnema quixensis and Phyzelaphryne miriamae (Amphibia: Anura; Leptodactylidae). Journal of Herpetology, 29(2): 304-307.

Cardoso, Adao J. and Heyer, W. Ronald 1995. Advertisement, aggressive, and possible seismic signals of the frog Leptodactylus syphax (Amphibia, Leptodactylidae). Alytes Paris, 13(2): 67-76.

Heyer, W. R., Donnelly, Maureen A., McDiarmid, Roy W., Hayek, Lee-Ann C. and Foster, Mercedes S. 1994. Essentials of Standardization and Quantification. In: Heyer, W. R., Measuring and Monitoring Biological Diversity: Standard Methods for Amphibians. Smithsonian Institution Press, pp.17-39.

Fellers, G. M., Drost, C. A. and Heyer, W. Ronald 1994. Handling Live Amphibians. In: Heyer, W. R., Measuring and Monitoring Biological Diversity: Standard Methods for Amphibians. Smithsonian Institution Press, pp.275-276.

Heyer, W. R., Donnelly, Maureen A., McDiarmid, Roy W., Hayek, Lee-Ann C. and Foster, Mercedes S. 1994. Conclusions and Recommendations. In: Heyer, W. R., Measuring and Monitoring Biological Diversity: Standard Methods for Amphibians. Smithsonian Institution Press, pp.271-273.

Jacobs, Jeremy F. and Heyer, W. Ronald 1994. Collecting Tissue for Biochemical Analysis. In: Heyer, W. R., Measuring and Monitoring Biological Diversity: Standard Methods for Amphibians. Smithsonian Institution Press, pp.299-301.

Heyer, W. R., Donnelly, Maureen A., McDiarmid, Roy W., Hayek, Lee-Ann C. and Foster, Mercedes S. 1994. Contents. In: Heyer, W. R., Measuring and Monitoring Biological Diversity: Standard Methods for Amphibians. Smithsonian Institution Press, pp.v-viii.

Heyer, W. R., Donnelly, Maureen A., McDiarmid, Roy W., Hayek, Lee-Ann C. and Foster, Mercedes S. 1994. Preface and Introduction. In: Heyer, W. R., Measuring and Monitoring Biological Diversity: Standard Methods for Amphibians. Smithsonian Institution Press, pp.xvii-3.

Heyer, W. R. 1994. Recording Frog Calls. In: Heyer, W. R., Measuring and Monitoring Biological Diversity: Standard Methods for Amphibians. Smithsonian Institution Press, pp.285-287.

Heyer, W. R., Donnelly, Maureen A., McDiarmid, Roy W., Hayek, Lee-Ann C. and Foster, Mercedes S. 1994. Measuring and monitoring biological diversity: standard methods for amphibians Smithsonian Institution Press, Washington and London. 364 pages.

Heyer, W. R. 1994. [Newsletter.]. Herpetologists' League Communications, 1(1): 4

Heyer, W. R. 1994. Hyla benitezi (Amphibia: Anura: Hylidae): first record for Brazil and its biogeographical significance. Journal of Herpetology, 28(4): 497-499.

Heyer, W. R. 1994. Prasenz epigaischer Raubarthropoden in Apfelanlagen - Ansatz zu einer Gefahrdungsbeurteilung. Nachrichtenblatt des Deutschen Pflanzenschutzdienstes Stuttgart, 46(10): 218-224.

Heyer, W. R. 1994. Variation within the Leptodactylus podicipinus-wagneri complex of frogs (Amphibia: Leptodactylidae). Smithsonian Contributions to Zoology, 546: i-124. doi:10.5479/si.00810282.546.i

Heyer, W. R., Donnelly, Maureen A. and McDiarmid, Roy W. 1994. Estimating Population Size. In: Heyer, W. R., Measuring and Monitoring Biological Diversity: Standard Methods for Amphibians. Smithsonian Institution Press, pp.183-205.

Heyer, W. R. 1994. Thread bobbins. In: Heyer, W. R., Donnelly, Maureen A., McDiarmid, Roy W., Hayek, Lee-Ann C. and Foster, Mercedes S., Measuring and Monitoring Biological Diversity: Standard Methods for Amphibians. Washington, D.C.: Smithsonian Institution Press, pp.153-155.

Heyer, W. R., Donnelly, Maureen A., McDiarmid, Roy W., Hayek, Lee-Ann C. and Foster, Mercedes S. 1994. Index. In: Heyer, W. R., Measuring and Monitoring Biological Diversity: Standard Methods for Amphibians. Smithsonian Institution Press, pp.351-359.

Heyer, W. R., Donnelly, Maureen A., McDiarmid, Roy W., Hayek, Lee-Ann C. and Foster, Mercedes S. 1994. Literature Cited. In: Heyer, W. R., Measuring and Monitoring Biological Diversity: Standard Methods for Amphibians. Smithsonian Institution Press, pp.329-350.

Heyer, W. R., Donnelly, Maureen A., McDiarmid, Roy W., Hayek, Lee-Ann C. and Foster, Mercedes S. 1994. Supplemental Approaches to Studying Amphibian Biodiversity. In: Heyer, W. R., Measuring and Monitoring Biological Diversity: Standard Methods for Amphibians. Smithsonian Institution Press, pp.143-182.

Heyer, W. R., Donnelly, Maureen A., McDiarmid, Roy W., Hayek, Lee-Ann C. and Foster, Mercedes S. 1994. Keys to a Successful Project. In: Heyer, W. R., Measuring and Monitoring Biological Diversity: Standard Methods for Amphibians. Smithsonian Institution Press, pp.41-73.

Heyer, W. R., Donnelly, Maureen A., McDiarmid, Roy W., Hayek, Lee-Ann C. and Foster, Mercedes S. 1994. Glossary. In: Heyer, W. R., Measuring and Monitoring Biological Diversity: Standard Methods for Amphibians. Smithsonian Institution Press, pp.325-327.

Heyer, W. R. and Perez, Linnette Garcia 1993. Description of the Advertisement Call and Resolution of the Systematic Status of Leptodactylus Gracilis Delattini Muller, 1968 (Amphibia, Leptodactylidae). Proceedings of the Biological Society of Washington, 106(1): 51-56.

Heyer, W. R., Chiang Lok, M. L. and Cruz, B. 1992. Zur Abundanzdynamik von Systena basalis Duval. und Andrector ruficornis (Oliv.) in Bohnenbestanden. Archiv fuer Phytopathologie und Pflanzenschutz, 28(1): 51-61.

Heyer, W. R. 1992. Zur Parasitierung der Getreidehahnchen Oulema spp. im Gebiet von Halle. Mitteilungen der Deutschen Gesellschaft fuer Allgemeine und Angewandte Entomologie, 8(1-3): 87-92.

Dubois, Alain and Heyer, W. Ronald 1992. Leptodactylus labialis, the Valid Name for the American Whitelipped Frog (Amphibia: Leptodactylidae). Copeia, 2: 584-585.

Heyer, W. R., Chiang Lok, M. L. and Cruz, B. 1991. Zur Fertilitat, Mortalitat und Altersstruktur der Popultionen von Empoasca kraemeri (Ross und Moore) in Bohnenbestanden (Phaseolus vulgaris L.) Kubas. Archiv fuer Phytopathologie und Pflanzenschutz, 27(3): 229-238.

Al Hussein, I. A., Heyer, W. Ronald and Wetzel, T. 1991. Untersuchungen zum Auftreten der Webespinnen (Araneae) in Winterweizen und ihre Beeinflussung durch ausgewahlte Insektizide. Archiv fuer Phytopathologie und Pflanzenschutz, 27(3): 219-228.

Heyer, W. R., Chiang Lok, M. L. and Cruz, B. 1991. Untersuchungen zur Entwicklungsdauer von Empoasca kraemeri (Ross und Moore) (Homoptera: Cicadellidae) und ihrem Massenwechsel in Bohnenbestanden (Phaseolus vulgaris L.) Kubas. Beitraege zur Tropischen Landwirtschaft und Veterinaermedizin, 29(4): 453-464.

Heyer, W. R. and Hardy, L. M. 1991. A New Species of Frog of the Eleutherodactylus-Lacrimosus Assembly from Amazonia, South-America (Amphibia, Anura, Leptodactylidae). Proceedings of the Biological Society of Washington, 104(3): 436-447.

Heyer, W. R. and Richter, S. 1990. Investigations into the temperature related development of the Serpentine leaf miner Liriomyza trifoli (Burgess) on beans (Phaseolus vulgaris L.). Beitraege zur Entomologie, 40(1): 259-264.

Heyer, W. R. and Wolf, A. J. 1989. Physalaemus-Crombiei (Amphibia, Leptodactylidae), a New Frog Species from Espirito-Santo, Brazil with Comments on the Physalaemus-Signifer Group. Proceedings of the Biological Society of Washington, 102(2): 500-506.

Heyer, W. R., Chiang Lok, M. L. and Cruz, B. 1989. Massenwechsel und Schadwirkung von Liriomyza trifolii (Burgess) in Bohnenbestanden Kubas. Archiv fuer Phytopathologie und Pflanzenschutz, 25(5): 487-496.

Heyer, W. R., Chiang Lok, M. L. and Cruz, B. 1989. Zur Populationsdynamik der Mottenschildlaus Bemisia tabaci Genn. in Bohnenbestanden der Republik Kuba. Archiv fuer Phytopathologie und Pflanzenschutz, 25(5): 473-479.

Heyer, W. R., Chiang Lok, M. L. and Cruz, B. 1989. Zur Populationsdynamik von Diabrotica balteata Lec. in Bohnenbestanden Kubas. Beitraege zur Entomologie, 39(1): 157-162.

Heyer, W. R. and Wetzel, T. 1989. Zum Einfluss der Populationsdichte auf die Nachkommenproduktion der Getreidehahnchen (Oulema spp.). Beitraege zur Entomologie, 39(2): 377-381.

Heyer, W. R. and Cruz, B. 1989. Kurze Mitteilung: Moglichkeit der Geschlechterdetermination beim Blattkafer Diabrotica balteata Lec. Beitraege zur Entomologie, 39(2): 391-392.

Heyer, W. R., Chiang Lok, M. L. and Cruz, B. 1989. Some aspects of biology and population dynamics of Diabrotica balteata Lec. and Systena basalis Duval. (Coleoptera, Chrysomelidae), maize and bean pests in Cuba. Acta Phytopathologica et Entomologica Hungarica, 24(1-2): 93-97.

Cocroft, Reginald B. and Heyer, W. Ronald 1988. Notes on the Frog Genus Thoropa (Amphibia: Leptodactylidae) with a Description of a New Species (Thoropa saxatilis). Proceedings of the Biological Society of Washington, 101(1): 209-220.

Heyer, W. R., Chiang Lok, M. L. and Cruz, B. 1988. Zum Geschlechterverhaltnis ausgewahlter Schadinsektenpopulationen in Bohnenbestanden. Beitraege zur Entomologie, 38(1): 189-194.

Heyer, W. R., Chiang Lok, M. L. and Cruz, B. 1988. Zum Einfluss der Temperatur und Wirtspflanze auf die Entwicklung von Andrector ruficornis (Oliv.) (Coleoptera: Chrysomelidae). Beitraege zur Entomologie, 38(1): 183-188.

Maxson, Linda R. and Heyer, W. Ronald 1988. Molecular systematics of the frog genus Leptodactylus (Amphibia: Leptodactylidae). Fieldiana Zoology, 41: 1-13.

Heyer, W. R., Rand, Austin Stanley, Dacruz, C. A. G. and Peixoto, O. L. 1988. Decimations, Extinctions, and Colonizations of Frog Populations in Southeast Brazil and their Evolutionary Implications. Biotropica, 20(3): 230-235.

Heyer, W. R. 1988. A Notable Collection of Cycloramphus (Amphibia, Leptodactylidae) from Bahia, Brazil, with a Description of a New Species (Cycloramphus-Migueli). Proceedings of the Biological Society of Washington, 101(1): 151-154.

Wassersug, Richard J. and Heyer, W. Ronald 1988. A survey of internal oral features of Leptodactyloid larvae (Amphibia: Anura). Smithsonian Contributions to Zoology, 457: 1-99. doi:10.5479/si.00810282.457

Heyer, W. R. and Vanzolini, P. E. 1988. Introduction and Summary of a Workshop on Neotropical Distribution Patterns held 12–16 January 1987. In: Vanzolini, P. E. and Heyer, W. Ronald, Proceedings of a Workshop on Neotropical Distribution Patterns Held 12–16 January 1987. Rio de Janeiro: Academia Brasileira de Ciências, pp.7-18.

Heyer, W. R. 1988. On Frog Distribution Patterns East of the Andes. In: Proceedings of a Workshop on Neotropical Distribution Patterns. Academia Brasileira de Ciéncias, pp.245-273.

Heyer, W. R. and Cocroft, R. B. 1986. Descriptions of two new species of Hylodes from the Atlantic forests of Brazil (Amphibia: Leptodactylidae). Proceedings of the Biological Society of Washington, 99(1): 100-109.

Heyer, W. R. 1985. Taxonomic and natural history notes on frogs of the genus Centrolenella (Amphibia: Centrolenidae) from southeastern Brasil and adjacent Argentina. Papeis Avulsos de Zoologia Sao Paulo, 36(1): 1-21.

Heyer, W. R. 1985. New species of frogs from Boraceia, Sao Paulo, Brazil. Proceedings of the Biological Society of Washington, 98(3): 657-671.

Vanzolini, P. E. and Heyer, W. Ronald 1985. The American herpetofauna and the interchange. Topics in Geobiology, 4: 475-487.

Heyer, W. R. 1984. The systematic status of Adenomera griseigularis Henle, with comments on systematic problems in the genus Adenomera (Amphibia: Leptodactylidae). Amphibia-Reptilia, 5(2): 97-100.

Heyer, W. R. 1984. Variation, systematics, and zoogeography of Eleutherodactylus guentheri and closely related species (Amphibia: Anura: Leptodactylidae). Smithsonian Contributions to Zoology, 402: 1-42. doi:10.5479/si.00810282.402

Heyer, W. R. 1984. Advances in Herpetology and Evolutionary Biology - Essays in Honor of Williams,ernest,e. - Rhodin,agj, Miyata,k. Science, 223(4632): 162

Crombie, Ronald I. and Heyer, W. Ronald 1983. Leptodactylus longirostris (Anura: Leptodactylidae): advertisement call, tadpole, ecological and distributional notes. Revista brasileira de biologia, 43(3): 291-296.

Heyer, W. R. and Pyburn, W. F. 1983. Leptodactylus riveroi, a new frog species from Amazonia, South America (Anura: Leptodactylidae). Proceedings of the Biological Society of Washington, 96(3): 560-566.

Heyer, W. R. 1983. Notes on the frog genus Cycloramphus (Amphibia: Leptodactylidae), with descriptions of two new species. Proceedings of the Biological Society of Washington, 96(3): 548-559.

Heyer, W. R. 1983. Variation and systematics of frogs of the genus Cycloramphus (Amphibia, Leptodactylidae). Arquivos de Zoologia Sao Paulo, 30(4): 235-339.

Wassersug, Richard J. and Heyer, W. Ronald 1983. Morphological correlates of subaerial existence in leptodactylid tadpoles associated with flowing water. Canadian Journal of Zoology, 61(4): 761-769.

Heyer, W. R. and Maxson, Linda R. 1983. Relationships, zoogeography and speciation mechanisms of frogs of the genus Cycloramphus (Amphibia, Leptodactylidae). Arquivos de Zoologia Sao Paulo, 30(5): 341-373.

Heyer, W. R. 1983. Clarification of the names Rana mystacea Spix, 1824, Leptodactylus amazonicus Heyer, 1978 and a description of a new species, Leptodactylus spixi (Amphibia: Leptodactylidae). Proceedings of the Biological Society of Washington, 96(2): 270-272.

Heyer, W. R. and Maxson, Linda R. 1982. Neotropical frog biogeography: paradigms and problems. American Zoologist, 22(2): 397-410.

Maxson, Linda R. and Heyer, W. Ronald 1982. Leptodactylid frogs and the Brasilian Shield: an old and continuing adaptive relationship. Biotropica, 14(1): 10-15.

Heyer, W. R. and Maxson, Linda R. 1982. Distributions, relationships and zoogeography of lowland frogs. The Leptodactylus complex in South America, with special reference to Amazonia. In: Prance, G. T., Biological diversification in the tropics. New York: Columbia University Press, pp.375-388.

Heyer, W. R., Daugherty, C. H. and Maxson, Linda R. 1982. Systematic resolution of the genera of the Crinia complex (Amphibia: Anura: Myobatrachidae). Proceedings of the Biological Society of Washington, 95(3): 423-427.

Heyer, W. R. 1982. Two new species of the frog genus Hylodes from Caparao, Minas Gerais, Brazil (Amphibia: Leptodactylidae). Proceedings of the Biological Society of Washington, 95(2): 377-385.

Drewry, George E., Heyer, W. Ronald and Rand, Austin Stanley 1982. A functional analysis of the complex call of the frog Physalaemus pustulosus. Copeia, 3: 636-645.

Maxson, Linda R., Heyer, W. Ronald and Maxson, R. D. 1981. Phylogenetic relationships of the Brasilian leptodactylid frog genera Craspedoglossa, Cycloramphus and Zachaenus. Experientia Basel, 37(2): 144-145.

Heyer, W. R. 1980. The calls and taxonomic positions of Hyla giesleri and Ololygon opalina (Amphibia: Anura: Hylidae). Proceedings of the Biological Society of Washington, 93(3): 655-661.

Heyer, W. R. and Crombie, Ronald I. 1979. Natural history notes on Craspedoglossa Stejnegeri and Thoropa petropolitana (Amphibia: Salientia, Leptodactylidae). Journal of the Washington Academy of Sciences, 69(1): 17-20.

Heyer, W. R. 1979. Annual variation in larva amphibian populations within a temperate pond. Journal of the Washington Academy of Sciences, 69(2): 65-74.

Heyer, W. R. and Mello, C. C. 1979. Descriptions of the advertising calls of Cycloramphus asper and Cycloramphus dubius (Amphibia: Leptodactylidae). Papeis Avulsos de Zoologia Sao Paulo, 32(15): 193-200.

Heyer, W. R. 1979. Systematics of the pentadactylus species group of the frog genus Leptodactylus (Amphibia, Leptodactylidae. Smithsonian Contributions to Zoology, 301: 1-43. doi:10.5479/si.00810282.301

Heyer, W. R. 1978. Systematics of the fuscus group of the frog genus Leptodactylus (Amphibia, Leptodactylidae). National History Museum of Los Angeles County, Scientific Bulletin, 8(29): 1-85.

Heyer, W. R. 1978. Variation in members of the Centrolenella eurygnatha complex (Amphibia: Centrolenidae) from Serra do Mar and Serra da Mantiqueira, Brasil. Papeis Avulsos de Zoologia Sao Paulo, 32(2): 15-33.

Heyer, W. R. 1978. [Review] "A Biogeographical Analysis of the Chihuahuan Desert through Its Herpetofauna.". Herpetological Review, 9(2): 50-51.

Heyer, W. R. and Muedeking, Miriam H. 1977. Notes on Tadpoles as Prey for Naiads and Turtles. Journal of the Washington Academy of Sciences, 66(4): 235-239.

Heyer, W. R. 1977. Foam Nest Construction in the Leptodactylid Frogs Leptodactylus pentadactylus and Physalaemus pustulosus (Amphibia, Anura, Leptodactylidae). Journal of Herpetology, 11(2): 225-228.

Heyer, W. R. 1977. Taxonomic Notes on Frogs from the Madeira and Purus Rivers, Brasil. Papeis Avulsus de Zoologia, 31(8): 141-162.

Heyer, W. R. 1977. A Discriminant Function Analysis of the Frogs of the Genus Adenomera (Amphibia, Leptodactylidae). Proceedings of the Biological Society of Washington, 89(51): 581-592.

Muedeking, Miriam H. and Heyer, W. Ronald 1976. Descriptions of the Eggs and Reproductive Patterns of Leptodactylus Pentadactylus (Amphibia: Leptodactylidae). Herpetologica, 32: 137-139.

Heyer, W. R. 1976. Notes on the Frog Fauna of the Amazon Basin. Acta Amazonica, 6(3): 369-378.

Heyer, W. R. and Muedeking, Miriam H. 1976. Notes on Tadpoles as Prey for Naiads and Turtles. Journal of Washington Academic Science, 66(4): 235-239.

Heyer, W. R. and Liem, David S. 1976. Analysis of the intergeneric relationships of the Australian frog family Myobatrachidae. Smithsonian Contributions to Zoology, 233: 1-29. doi:10.5479/si.00810282.233

Heyer, W. R. 1976. Studies in larval amphibian habitat partitioning. Smithsonian Contributions to Zoology, 242: 1-27. doi:10.5479/si.00810282.242

Straughan, Ian R. and Heyer, W. Ronald 1976. A Functional Analysis of the Mating Calls of the Neotropical Frog Genera of the Leptodactylus Complex (Amphibia, Leptodactylidae). Papeis Avulsos de Zoologia, 29(23): 221-245.

Heyer, W. R. 1976. [Review] "Evolution of the Desert Biota.". Copeia, 3: 614-615.

Heyer, W. R. 1976. The Presumed Tadpole of Paratelmatobius lutzi (Amphibia, Leptodactylidae). Papéis Avulsos, Sāo Paulo, 30(10): 133-135.

Heyer, W. R. 1975. Adenomera lutzi (Amphibia: Leptodactylidae), a New Species of Frog from Guyana. Proceedings of the Biological Society of Washington, 88(28): 315-318.

Pyburn, W. F. and Heyer, W. Ronald 1975. Identity and Call of Frog, Leptodactylus-Stenodema. Copeia, 3: 585-587.

Heyer, W. R. 1975. A preliminary analysis of the intergeneric relationships of the frog family Leptodactylidae. Smithsonian Contributions to Zoology, 199: 1-55. doi:10.5479/si.00810282.199

Heyer, W. R., McDiarmid, Roy W. and Weigmann, D. L. 1975. Tadpoles, predation and pond habitats in the tropics. Biotropica, : 100-111.

Heyer, W. R. 1974. Relationships of the Marmoratus Species Group (Amphibia, Leptodactylidae) Within the Subfamily Leptodactylinae. Contributions in Science: Natural History Museum, Los Angeles County, 253: 1-45.

Heyer, W. R. 1974. Vanzolinius, A New Genus Proposed for Leptodactylus Discodactylus (Amphibia, Leptodactylidae). Proceedings of the Biological Society of Washington, 87(11): 81-90.

Heyer, W. R. and Diment, Judith M. 1974. The Karyotype of Vanzolinius Discodactylus and Comments on Usefulness of Karyotypes in Determining Relationships in the Leptodactylus-Complex (Amphibia, Leptodactylidae). Proceedings of the Biological Society of Washington, 87(29): 327-336.

Heyer, W. R. 1974. Niche Measurements of Frog Larvae from a Seasonal Tropical Location in Thailand. Ecology, 55(3): 651-656.

McDiarmid, Roy W., Heyer, W. Ronald and Weigman, Diana L. 1974. Tadpoles, predation and pond size in the tropics. Herpetological Review, 5(3): 76

Heyer, W. R. and Berven, Keith A. 1973. Species diversities of herpetofaunal samples from similar microhabitats at two tropical sites. Ecology, 54(3): 642-645.

Heyer, W. R. and Bellin, Melinda S. 1973. Ecological Notes on Five Sympatric Leptodactylus (Amphibia, Leptodactylidae) from Ecuador. Herpetologica, 29(1): 66-72.

Heyer, W. R. 1973. Ecological Interactions of Frog Larvae at a Seasonal Tropical Location in Thailand. Journal of Herpetology, 7(4): 337-361.

Heyer, W. R. 1973. Systematics of the Marmoratus Group of the Frog Genus Leptodactylus (Amphibia, Leptodactylidae). Contributions in Science, 251: 1-50.

Heyer, W. R. 1972. Leptodactylus labialis. 104. , : 000

Heyer, W. R. 1971. The tadpole of Rana pileata. Herpetological Review, 3: 95

Heyer, W. R. and Peters, James Arthur 1971. The frog genus Leptodactylus in Ecuador. Proceedings of the Biological Society of Washington, 84: 163-170.

Heyer, W. R. 1971. Descriptions of some tadpoles from Thailand. Fieldiana Zoology, 58: 83-91.

Heyer, W. R. 1971. Mating calls of some frogs from Thailand. Fieldiana Zoology, 58: 61-82.

Heyer, W. R. 1970. Studies on the frogs of the genus Leptodactylus (Amphibia: Leptodactylidae). 6. Biosystematics of the melanonotus group. Contr.Sci., No.,191: 1-48.

Heyer, W. R. 1970. Gliding speeds of Ptychozoon lionatum (Reptilia: Gekkonidae) and Chrysopelea ornata (Reptilia: Colubridae). Herpetologica, 26: 317-319.

Heyer, W. R. 1969. Biosystematic studies on the frog genus Leptodactylus. Dissertation Abstracts, 29B: 3970-3971.

Heyer, W. R. 1969. The adaptive ecology of the species groups of the genus Leptodactylus (Amphibia, Leptodactylidae). Evolution, 23: 421-428.

Heyer, W. R. and Silverstone, P. A. 1969. The larva of the frog Leptodactylus hylaedactylus (Leptodactylidae). Fieldiana Zoology, 51: 141-145.

Heyer, W. R. 1969. Studies on frogs of the genus Leptodactylus (Amphibia, Leptodactylidae). 5. Taxonomic notes on L. latinasus, Rhodonotus, Romani and Wuchereri. Herpetologica, 25: 1-8.

Heyer, W. R. 1969. Studies on the genus Lepodactylus (Amphibia, Leptodactylidae). 3. A redefinition of the genus Leptodactylus and a description of a new genus of Leptodactylid frogs. Contr.Sci., 155: 1-14.

Dixon, J. R. and Heyer, W. Ronald 1968. Anuran succession in a temporary pond in Colima, Mexico. Bulletin of the Southern California Academy of Sciences, 67: 129-137.

Heyer, W. R. 1968. The proper name for the type-species of the genus Leptodactylus. Copeia, 1968: 160-162.

Heyer, W. R. 1968. Studies on the genus Leptodactylus (Amphibia:Leptodactylidae). 2. Diagnosis and distribution of the Leptodactylus of Costa Rica. Revta. Revista De Biología Tropical, 16: 171

Heyer, W. R. 1967. A herpetofaunal study of an ecological transect through the Cordillera de Tilaran, Costa Rica. Copeia, 1967: 259-271.

Savage, Jay M. and Heyer, W. Ronald 1967. Variation and distribution in the tree-frog genus Phyllomedusa in Costa Rica, central America. Beiträge zur Neotropischen Fauna, 5: 111-131.

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