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Smithsonian National Museum of Natural History

Department of Vertebrate Zoology

Division of Amphibians & Reptiles

Phyllomedusa vaillantii
Phyllomedusa vaillantii Ecuador, Pastaza Province. Photographed by William W. Lamar
bar George Zug
    George R. Zug
    Curator Emeritus

  • Phone: (202) 633-0738
  • Fax: (202) 633-0182

  • Mailing Address:
    Smithsonian Institution
    PO Box 37012, MRC 162
    Washington, DC 20013-7012

  • Shipping Address:
    Smithsonian Institution
    National Museum of Natural History
    10th and Constitution Ave, NW
    Washington, DC 20560-0162



B.S., Albright College, 1960
M.S., University of Florida, 1963
Ph.D., University of Michigan, 1968

Research Interests

Herpetology/biology and systematics of amphibians and reptiles; currently focused on the systematics of the herpetofauna of Oceania and Burma.

Recent Publications

Zug, George R. and Kaiser, Hinrich 2014. A new species of four-toed skink (Squamata: Scincidae: Carlia peronii species group) from Pulau Sukur, Indonesia, and biogeographic notes on the herpetofauna of Flores and Komodo. Proceedings of the Biological Society of Washington, 126(4): 379-392. doi:10.2988/0006-324X-126.4.379

Mulcahy, Daniel G., Martínez-Gómez, Juan E., Aguirre-León, Gustavo, Cervantes-Pasqualli, Juan and Zug, George R. 2014. Rediscovery of an Endemic Vertebrate from the Remote Islas Revillagigedo in the Eastern Pacific Ocean: The Clarión Nightsnake Lost and Found. Plos One, 9(5): 1-8. doi:10.1371/journal.pone.0097682

Nguyen, Truong Q., Botov, Andreas, Le, Minh Duc, Nophaseud, Liphone, Zug, George R., Bonkowski, Michael and Ziegler, Thomas 2014. A new species of Hemiphyllodactylus (Reptilia: Gekkonidae) from northern Laos. Zootaxa, 3827(1): 45-56.

Böhm, Monika, Collen, Ben, Baillie, Jonathan E. M., Bowles, Philip, Chanson, Janice, Cox, Neil, Hammerson, Geoffrey, Hoffmann, Michael, Livingstone, Suzanne R., Ram, Mala, Rhodin, Anders G. J., Stuart, Simon N., van Dijk, Peter Paul, Young, Bruce E., Afuang, Leticia E., Aghasyan, Aram, García, Andrés, Aguilar, Cé, Ajtic, Rastko, Akarsu, Ferdi, Alencar, Laura R. V., Allison, Allen, Ananjeva, Natalia, Anderson, Steve, Andrén, Claes, et al 2013. The conservation status of the world's reptiles. Biological Conservation, 157: 372-385. doi:10.1016/j.biocon.2012.07.015

Zug, George R. 2013. Literature resources For "Reptiles And Amphibians Of The Pacific Islands: a comprehensive guide". Smithsonian Herpetological Information Service, 143: 1-32.

Fisher, Robert N., Uili, Moeumu, Enoka, Fialelei and Zug, George R. 2013. Clarifying the distributional records for Hemiphyllodactylus typus (Bleeker, 1860) from Central Oceania. Herpetology Notes, 6: 247-250.

Zug, George R. and Kaiser, Hinrich 2013. Comment on the proposed conservation of Kalophrynus Tschudi, 1838 (Amphibia, Anura, Microhylidae) by designation of a neotype for its type species Kalophrynus pleurostigma Tschudi, 1838. Bulletin of Zoological Nomenclature, 71(1): 28

Zug, George R. 2013. Reptiles and Amphibians of the Pacific Islands: a comprehensive guide Berkeley: University of California Press. 306 pages.

Zug, George R. and Kaiser, Hinrich 2013. Kalophrynus Tschudi, 1838 (Amphibia, Anura, Microhylidae): proposed conservation by designation of a neotype for its type species Kalophrynus pleurostigma Tschudi, 1838. Bulletin of Zoological Nomenclature, 70(2): 86-88.

Zug, George R. and Fisher, R. N. 2012. A preliminary assessment of the Nactus pelagicus species group (Squamata: Gekkonidae) in New Guinea and a new species from the Admiralty Islands. Zootaxa, 3257: 22-37.

Zug, George R. 2012. Lizards of Tonga with a description of a new Tongan treeskink (Squamata: Scincidae: Emoia samoensis Group). Pacific Science, 66(2): 225-237. doi:10.2984/66.2.9

Zug, George R. 2012. A new species of treeskink (Squamata: Scincidae: Emoia samoensis species group) from Rotuma, south-central Pacific. Proceedings of the Biological Society of Washington, 125(1): 74-84. doi:10.2988/11-24.1

Zug, George R.. Data resources for Tropical Asian Dry Forest Amphibians and Reptiles. Smithsonian Institution. 2012.

Austin, Christopher C., Rittmeyer, Eric N., Oliver, Lauren A., Andermann, John O., Zug, George R., Rodda, Gordon H. and Jackson, Nathan D. 2011. The bioinvasion of Guam: inferring geographic origin, pace, pattern and process of an invasive lizard (Carlia) in the Pacific using multi-locus genomic data. Biological Invasions, 13(9): 1951-1967. doi:10.1007/s10530-011-0014-y

Zug, George R. 2011. Tropical Asian Dry Forest Amphibians and Reptiles: A regional comparison of ecological communities. In: McShea, William J., Davies, Stuart James and Bjumpakpjan, Naris, The Ecology and Conservation of Seasonally Dry Forest in Asia. Washington, DC: Smithsonian Institution Scholarly Press, pp.275-303.

Zug, George R., Gotte, Steve W. and Jacobs, Jeremy F. 2011. Pythons in Burma: Short-tailed Pythons (Reptilia: Squamata). Proceedings of the Biological Society of Washington, 124(2): 112-136. doi:10.2988/10-34.1

Zug, George R., Hamilton, Alison M. and Austin, Christopher C. 2011. A new Emoia samoensis group lizard (Squamata: Scincidae) from the Cook Islands, South-central Pacific. Zootaxa, 2765: 47-57.

Zug, George R. 2010. An Outlying Carlia Population from Java and Comments on Species Groups within the Genus Carlia (Reptilia: Squamata: Scincidae). Proceedings of the California Academy of Sciences, 61(8): 389-408.

Zug, George R., Jacobs, Jeremy F., Vindum, Jens V. and Win, Kyi Soe 2010. Non-reproductive seasonal colour change in a population of Calotes "versicolor" from Myanmar (Squamata: Agamidae). Salamandra, 46(2): 104-107.

Austin, Christopher C., Rittmeyer, Eric N., Richards, Stephen J. and Zug, George R. 2010. Molecular phylogeny, historical biogeography and body size evolution in Pacific crocodile skinks Tribolonotus (Squamata; Scincidae). Molecular Phylogenetics and Evolution, 57: 227-236.

Balmer, Oliver, Ciofi, Claudio, Galbraith, David A., Swingland, Ian R., Zug, George R. and Caccone, Adalgisa 2010. Population genetic structure of Aldabra giant tortoises. Journal of Heredity, 102(1): 29-37. doi:10.1093/jhered/esq096

Furness, Andrew I., McDiarmid, Roy W., Heyer, W. Ronald and Zug, George R. 2010. Oviduct modifications in foam-nesting frogs, with emphasis on the genus Leptodactylus (Amphibia, Leptodactylidae). South American Journal of Herpetology, 5(1): 13-29. doi:10.2994/057.005.0102

Watling, Dick, Wynn, Addison H. and Zug, George R. 2010. Rediscovery of the Taveuni blind snake. Oryx, 44(2): 165-166.

Zug, George R. 2010. Speciation and dispersal in a low diversity taxon: the Slender Geckos Hemiphyllodactylus (Reptilia, Gekkonidae). Smithsonian Contributions to Zoology, 631: 1-70. doi:10.5479/si.00810282.631

Hamilton, Alison M., Zug, George R. and Austin, Christopher C. 2010. Biogeographic anomaly or human introduction: a tree skink (Reptilia: Squamata) in the Cook Islands?. Biological Journal of the Linnean Society, 100: 318-328.

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