Education

B.A., M.A., Pacific Lutheran University.
Ph.D., Catholic University.

Responsibilities

Collaborates on research of the Neotropical frog genus Leptodactylus.  Has primary responsibility for creation and maintenance of the Leptodactylus EndNote® bibliography (http://learning.richmond.edu/Leptodactylus/bibliography.html).

Selected Publications

Muedeking (Heyer), M. H. and W. R. Heyer.  1976.  Descriptions of eggs and reproductive patterns of Leptodactylus pentadactylus (Amphibia: Leptodactylidae).  Herpetologica 32(2):137-139.

Heyer, W. R. and M. M.Heyer.  2002.  Leptodactylus elenae.  Catalogue of American Amphibians and Reptiles (742):1-5

Heyer, M. M., W. R. Heyer, S. Spear, and R. O. de Sá.  2003.  Leptodactylus mystacinus.  Catalogue of American Amphibians and Reptiles (767):1-11.

Heyer, W.  R., and M. M. Heyer.  2005.  Leptodactylus knudseni.  Catalogue of American Amphibians and Reptiles (807):1-12

Heyer, M. M., W. R. Heyer, and R. O. de Sá.  2006.  Leptodactylus fragilis.  Catalogue of American Amphibians and Reptiles (830):1-26.