Education

B.A., M.A., Pacific Lutheran University.
Ph.D., Catholic University.

Responsibilities

Collaborates on research of the Neotropical frog genus Leptodactylus. Has primary responsibility for creation and maintenance of the Leptodactylinae EndNote® Bibliography.

Recent Publications

Heyer, W. R. and Heyer, Miriam M. 2012. Systematics, distribution, and bibliography of the frog Leptodactylus validus Garman, 1888 (Amphibia: Leptodactylidae). Proceedings of the Biological Society of Washington, 125(3): 276-294. doi:10.2988/12-04.1

Heyer, Miriam M., Heyer, W. R. and de Sa, Rafael O. 2012. Leptodactylus pentadactylus. Catalogue of American Amphibians and Reptiles, 887: 1-48.

Heyer, W. R., Heyer, Miriam M. and De Sá, Rafael Omar 2010. Leptodactylus syphax. Catalogue of American Amphibians and Reptiles, 868: 1-9.

Heyer, W. R., Heyer, Miriam M. and De Sá, Rafael Omar 2010. Leptodactylus savagei. Catalogue of American Amphibians and Reptiles, 867: 1-19.

Heyer, Miriam H., Heyer, W. Ronald and De Sá, Rafael Omar 2009. Bibliography of the frogs of the Leptodactylus clade - Adenomera, Hydrolaetare, Leptodactylus, Lithodytes (Amphibia, Anura, Leptodactylidae). Volume 2. Indices. Smithsonian Herpetological Information Service, 2: 1-167.

Heyer, Miriam H., Heyer, W. Ronald and De Sá, Rafael Omar 2009. Bibliography of the frogs of the Leptodactylus clade - Adenomera, Hydrolaetare, Leptodactylus, Lithodytes (Ampbibia, Anura, Leptodactylidae). Volume 1. References. Smithsonian Herpetological Information Service, 1: 1-677.