Related Links

Frogs of Boracéia in PDF Format (Online Audio Supplement)
Information Bank For Leptodactylus

Education

Ph.D.,University of Southern California, 1968, Biology.
M.S., University of Southern California, 1965, Biology.
B.A., Pacific Lutheran University, 1963, Biology.

Research Interests

Herpetology, Systematics, Evolution, Ecology – current focus is on systematics and biogeography of the Neotropical frog genus Leptodactylus.

Selected Publications

2005. Heyer, W. R.  Variation and taxonomic clarification of the large species of the Leptodactylus pentadactylus species group (Amphibia: Leptodactylidae) from Middle America, northern  South America, and Amazonia.  Arquivos de Zoologia, 37(3):269-348.

2005. Hayek, L. C., & W. R. Heyer.  Determining sexual dimorphism in frog measurement data:  Integration of statistical significance, measurement error, effect size and biological significance.  Anais da Academia Brasileira de Ciências, 77(1):45-76.

2001. Wynn, A., & W. R. Heyer.  Do geographically widespread species of tropical amphibians exist?  An estimate of genetic relatedness within the Neotropical frog Leptodactylus fuscus (Schneider 1799) (Anura Leptodactylidae).  Tropical Zoology, 14:255-285.

1994. Heyer, W. R.  Variation within the Leptodactylus podicipinus-wagneri complex of frogs (Amphibia:  Leptodactylidae).  Smithsonian Contributions to Zoology, (546):1-124. [PDF]

1990. Heyer. W. R., A. S. Rand, C. A. G. Cruz, O. L. Peixoto, & C. E. Nelson.  Frogs of Boracéia.  Arquivos de Zoologia, 31:231-410.

1983.  Heyer, W. R. Variation and systematics of frogs of the genus Cycloramphus (Amphibia, Leptodactylidae).  Arquivos de Zoologia, 30:235-339.

1983.  Heyer, W. R., & L. R. Maxson.  Relationships, zoogeography, and speciation mechanisms of frogs of the genus Cycloramphus (Amphibia, Leptodactylidae).  Arquivos de Zoologia, 30:341-373.

1978.  Heyer, W. R.  Systematics of the fuscus group of the frog genus Leptodactylus (Amphibia, Leptodactylidae).  Natural History Museum of Los Angeles County Science Bulletin, (29):1-85.

1975.  Heyer, W. R., R. W. McDiarmid, & D. L. Weigmann.  Tadpoles, predation and pond habitats in the tropics.  Biotropica, 7:100-111.

1975.  Heyer, W. R.  A preliminary analysis of the intergeneric relationships of the frog family Leptodactylidae.  Smithsonian Contributions to Zoology, (199):1-55.

Complete List of Heyer's Publications and Downloadable PDFs