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Smithsonian National Museum of Natural History
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Department of Vertebrate Zoology

Division of Amphibians & Reptiles

Phyllomedusa vaillantii
Phyllomedusa vaillantii Ecuador, Pastaza Province. Photographed by William W. Lamar
bar Ronald Heyer
    W. Ronald Heyer
    Research Zoologist

  • Phone: (202) 633-0730
  • Fax: (202) 357-2986
  • E-mail: heyerr@si.edu

  • Mailing Address:
    Smithsonian Institution
    PO Box 37012, MRC 162
    Washington, DC 20013-7012

  • Shipping Address:
    Smithsonian Institution
    National Museum of Natural History
    10th & Constitution Ave NW
    Washington, DC 20560-0162

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Related Links

Complete List of Heyer's Publications and Downloadable PDFs
Frogs of Boracéia in PDF Format (Online Audio Supplement)
Leptodactylinae EndNote® Bibliography

Education

Ph.D.,University of Southern California, 1968, Biology.
M.S., University of Southern California, 1965, Biology.
B.A., Pacific Lutheran University, 1963, Biology.

Research Interests

Herpetology, Systematics, Evolution, Ecology; current focus is on systematics and biogeography of the Neotropical frog genus Leptodactylus.

Recent Publications

Heyer, W. R. 2014. Morphological analyses of frogs of the Leptodactylus latrans complex (Amphibia, Leptodactylidae) from selected localities in South America. Proceedings of the Biological Society of Washington, 126(4): 369-378. doi:10.2988/0006-324X-126.4.369

Heyer, W. R. 2014. Morphological analyses of frogs of the Leptodactylus latrans complex (Amphibia, Leptodactylidae) from selected localities in South America. Proceedings of the Biological Society of Washington, 126(4): 369-378.

Heyer, W. R. and Heyer, Miriam M. 2013. Systematics, distribution, and bibliography of the frog Leptodactylus insularum Barbour, 1906 (Amphibia: Leptodactylidae). Proceedings of the Biological Society of Washington, 126(3): 204-233. doi:10.2988/0006-324X-126.3.204

Heyer, Miriam M., Heyer, W. R. and de Sa, Rafael O. 2012. Leptodactylus pentadactylus. Catalogue of American Amphibians and Reptiles, 887: 1-48.

Heyer, W. R. and Heyer, Miriam M. 2012. Systematics, distribution, and bibliography of the frog Leptodactylus validus Garman, 1888 (Amphibia: Leptodactylidae). Proceedings of the Biological Society of Washington, 125(3): 276-294. doi:10.2988/12-04.1

Heyer, W. R. and de Sá, Rafael O. 2011. Variation, Systematics, and Relationships of the Leptodactylus bolivianus Complex (Amphibia: Anura: Leptodactylidae). Smithsonian Contributions to Zoology, 635: 1-58. doi:10.5479/si.00810282.635.1

Laura Ponssa, Maria, Heyer, W. Ronald and De Sá, Rafael Omar 2010. Phylogeny of the genus Leptodactylus (Anura, Leptodactylidae) based on characters of morphology and mode of life. Cladistics, 26(2): 222

Heyer, W. R., Heyer, Miriam M. and De Sá, Rafael Omar 2010. Leptodactylus syphax. Catalogue of American Amphibians and Reptiles, 868: 1-9.

Heyer, W. R., Heyer, Miriam M. and De Sá, Rafael Omar 2010. Leptodactylus savagei. Catalogue of American Amphibians and Reptiles, 867: 1-19.

Lavilla, Esteben O., Langone, Jose A., Caramaschi, Ulisses, Heyer, W. Ronald and De Sá, Rafael Omar 2010. The identification of Rana ocellata Linnaeus, 1758. Nomenclatural impact on the species currently known as Leptodactylus ocellatus (Leptodactylidae) and Osteopilus brunneus (Gosse, 1851)(Hylidae). Zootaxa, 2346: 1-16.

Furness, Andrew I., McDiarmid, Roy W., Heyer, W. Ronald and Zug, George R. 2010. Oviduct modifications in foam-nesting frogs, with emphasis on the genus Leptodactylus (Amphibia, Leptodactylidae). South American Journal of Herpetology, 5(1): 13-29. doi:10.2994/057.005.0102

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