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Department of Vertebrate Zoology

Division of Amphibians & Reptiles

Phyllomedusa vaillantii
Phyllomedusa vaillantii Ecuador, Pastaza Province. Photographed by William W. Lamar
bar Ronald Heyer
    W. Ronald Heyer
    Research Zoologist

  • Phone: (202) 633-0730
  • Fax: (202) 357-2986
  • E-mail:

  • Mailing Address:
    Smithsonian Institution
    PO Box 37012, MRC 162
    Washington, DC 20013-7012

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    Smithsonian Institution
    National Museum of Natural History
    10th & Constitution Ave NW
    Washington, DC 20560-0162


Related Links

Complete List of Heyer's Publications and Downloadable PDFs
Frogs of Boracéia in PDF Format (Online Audio Supplement)
Leptodactylinae EndNote® Bibliography


Ph.D.,University of Southern California, 1968, Biology.
M.S., University of Southern California, 1965, Biology.
B.A., Pacific Lutheran University, 1963, Biology.

Research Interests

Herpetology, Systematics, Evolution, Ecology; current focus is on systematics and biogeography of the Neotropical frog genus Leptodactylus.

Recent Publications

Heyer, W. Ronald and Heyer, Miriam M. 2016. Biographical sketch and bibliography of W. Ronald Heyer. Smithsonian Herpetological Information Service, 147: 1-16. doi:10.5479/si.23317515.147.1

de Sá, Rafael O., Grant, Taran, Camargo, Arley, Heyer, W. Ronald, Ponssa, Maria L. and Stanley, Edward 2014. Systematics of the Neotropical Genus Leptodactylus Fitzinger, 1826 (Anura: Leptodactylidae): Phylogeny, the Relevance of Non-molecular Evidence, and Species Accounts. South American Journal of Herpetology, 9: S1-S128. doi:10.2994/SAJH-D-13-00022.1

Heyer, W. R. 2014. Morphological analyses of frogs of the Leptodactylus latrans complex (Amphibia, Leptodactylidae) from selected localities in South America. Proceedings of the Biological Society of Washington, 126(4): 369-378. doi:10.2988/0006-324X-126.4.369

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