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Department ofVertebrate Zoology

Division of Fishes

Barracuda
Carl Hansen
© Smithsonian Institution
bar Rich Vari
    Richard Vari
    Interim Associate Director for Science, Curator of Fishes

  • Phone: (202)633-1207
  • Fax: (202)633-0182
  • E-mail: varir[at]si.edu

  • Mailing Address:
    Smithsonian Institution
    PO Box 37012, MRC 159
    Washington, DC 20013-7012

  • Shipping Address:
    Smithsonian Institution
    National Museum of Natural History
    10th and Constitution Ave, NW
    Washington, DC 20560-0159
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Education

Ph.D. 1976, City University of New York
B.S. 1971, New York University

Research Interests

Taxonomy, phylogeny and biogeography of South America, African, and South Asian freshwater fishes, in particular characiforms and siluriforms.

Recent Publications

Melo, Bruno F., Ochoa, Luz E., Vari, Richard P. and Oliveira, Claudio 2016. Cryptic species in the Neotropical fish genus Curimatopsis (Teleostei, Characiformes). Zoologica Scripta, doi:10.1111/zsc.12178

Melo, Bruno F., Sidlauskas, Brian L., Hoekzema, Kendra, Frable, Benjamin W., Vari, Richard P. and Oliveira, Claudio 2016. Molecular phylogenetics of the Neotropical fish family Prochilodontidae (Teleostei: Characiformes). Molecular phylogenetics and evolution, 102: 189-201. doi:10.1016/j.ympev.2016.05.037

Dillman, Casey B., Sidlauskas, Brian L. and Vari, Richard P. 2016. A morphological supermatrix-based phylogeny for the Neotropical fish superfamily Anostomoidea (Ostariophysi: Characiformes): phylogeny, missing data and homoplasy. Cladistics, 32(3): 276-296. doi:10.1111/cla.12127

Dutra, Guilherme Moreira, Jerep, Fernando C., Vari, Richard P. and de Santana, Carlos David 2015. The pseudotympanum in the Gymnotiformes (Teleostei, Ostariophysi, Otophysi): homology and evolution of a previously unexplored system in Neotropical electric fishes. Zoological Journal of the Linnean Society, 174(1): 114-129. doi:10.1111/zoj.12221

Carvalho, Murilo and Vari, Richard P. 2015. Development of the splanchnocranium in Prochilodus argenteus (Teleostei: Characiformes) with a discussion of the basal developmental patterns in the Otophysi. Zoology, 118(1): 34-50. doi:10.1016/j.zool.2014.10.002

García-Roselló, Emilio, Guisande, Cá, Manjarrés-Hernández, Ana, González-Dacosta, Jacinto, Heine, Juergen, Pelayo-Villamil, Patricia, González-Vilas, Luis, Granado-Lorencio, Carlos, Lobo, Jorge M., Vaamonde, Antonio and Vari, Richard P. 2015. Can we derive macroecological patterns from primary Global Biodiversity Information Facility data?. Global Ecology and Biogeography, 24(3): 335-347. doi:10.1111/geb.12260

Pelayo-Villamil, Patricia, Guisande, Cá, Vari, Richard P., Manjarrés-Hernández, Ana, García-Roselló, Emilio, González-Dacosta, Jacinto, Heine, Jü, González Vilas, Luis, Patti, Bernardo, Quinci, Enza María, Jiménez, Luz Fernanda, Granado-Lorencio, Carlos, Tedesco, Pablo A. and Lobo, Jorge M. 2015. Global diversity patterns of freshwater fishes – potential victims of their own success. Diversity and Distributions, 21(3): 345-356. doi:10.1111/ddi.12271

Vari, Richard P. and Calegari, Barbara B. 2014. New species of the catfish genus Tatia (Siluriformes:Auchenipteridae) from the rio Teles Pires, upper rio Tapajós basin, Brazil. Neotropical Ichthyology, 12(4): 667-674. doi:10.1590/1982-0224-20130193

Calegari, Barbara B., Reis, Roberto E. and Vari, Richard P. 2014. Miniature catfishes of the genus Gelanoglanis (Siluriformes: Auchenipteridae): monophyly and the description of a new species from the upper rio Tapajós basin, Brazil. Neotropical Ichthyology, 12(4): 699-706. doi:10.1590/1982-0224-20130233

Fernández, L., Sanabria, E. A., Quiroga, L. B. and Vari, Richard P. 2014. A new species of Silvinichthys (Siluriformes, Trichomycteridae) lacking pelvic fins from mid-elevation localities of the southern Andes, with comments on the genus. Journal of Fish Biology, 84(2): 372-382. doi:10.1111/jfb.12291

Jerep, Fernando C. and Vari, Richard P. 2014. New Species of Nannocharax (Characiformes: Distichodontidae) from the Northwestern Congo River Basin, with an Appraisal of Hemigrammocharax. Copeia, 1: 44-49. doi:10.1643/CI-13-048

Jerep, Fernando C., Vari, Richard P. and Vreven, Emmanuel 2014. Nannocharax dageti, a new distichodontid from the Democractic Republic of the Congo and Zambia (Teleostei: Characiformes). Ichthyological Exploration of Freshwaters, 24(4): 361-369.

Bruno, F. M. and Vari, Richard P. 2014. New species of Cyphocharax (Characiformes: Curimatidae) from the upper rio Negro, Amazon basin. Neotropical Ichthyology, 12(2): 327-332. doi:10.1590/1982-0224-20130153

Melo, Bruno F., Sidlauskas, Brian L., Hoekzema, Kendra, Vari, Richard P. and Oliveira, Claudio 2014. The first molecular phylogeny of Chilodontidae (Teleostei: Ostariophysi: Characiformes) reveals cryptic biodiversity and taxonomic uncertainty. Molecular Phylogenetics and Evolution, 70: 286-295. doi:10.1016/j.ympev.2013.09.025

Datovo, Alessio and Vari, Richard P. 2014. The adductor mandibulae muscle complex in lower teleostean fishes (Osteichthyes: Actginopterygii):comparative anatomy, synonymy, and phylogenetic implications. Zoological Journal of the Linnean Society, 171(3): 554-622. doi:10.1111/zoj.12142

Harrison, Ian, Evans, Malcolm, Lowe-McConnell, Rosemary, Parenti, Lynne R., Sanford, Chris, Snyder, Lex, Stiassny, Melanie and Vari, Richard P. 2013. Gordon John Howes 1938–2013 (Fish Systematist). Journal of fish biology, 83(5): 1085-1093. doi:10.1111/jfb.12260

de Queiroz, Luiz J., Torrente-Vilara, Gislene, Barros, Bruno S. F. and Vari, Richard P. 2013. Familia Ctenoluciidae. In: de Queiroz, Luiz J., Torrente-Vilara, Gislene, Ohara, Willian M., Silva Pires, R. H., Zuanon, J. and Costa Doria, C. R., Peixes do Rio Madeira, Volume 2. Sao Paulo: Santo Antonio, pp.91-95.

de Queiroz, Luis J., Ohara, Willian M. and Vari, Richard P. 2013. Familia Prochilodontidae. In: de Qieiroz, Luis J., Torrente-Vilara, G., Ohara, Willian M., Silva Pires, R. H., Zuanon, J. and Costa Doria, C. R., Peixes do Rio Madeira, Volume 1. Sao Paulo: Santo Antonio, pp.140-145.

Torrenti-Vilara, Gislene, de Queiroz, Luiz and Vari, Richard P. 2013. Familia Chilodontidae. In: de Queiroz, Luiz, Torrente-Vilara, Gislene, Ohara, Willian M., Silva Pires, R. H., Zuanon, J. and and Costa Doria, C. R., Peixes do Rio Madeira, Volume 1. Sao Paulo: Santo Antonio, pp.166-171.

Vari, Richard P. and Ferraris, Carl J. 2013. Two New Species of the Catfish Genus Tatia (Siluriformes: Auchenipteridae) from the Guiana Shield and a Reevaluation of the Limits of the Genus. Copeia, 2013(3): 396-402. doi:10.1643/CI-12-115

Datovo, Alessio and Vari, Richard P. 2013. The jaw adductor muscle complex in teleostean fishes: evolution, homologies and revised nomenclature (Osteichthyes: Actinopterygii). PLoS ONE, 8(4): 1-23. doi:10.1371/journal.pone.0060846

De Santana, Carlos and Vari, Richard P. 2013. Brown ghost electric fishes of the Apteronotus leptrorhynchus species-group (Ostariophysi, Gymnnotiformes); monophyly, major clades, and revision. Zoological Journal of the Linnean Society, 168(3): 564-596. doi:10.1111/zoj.12022

Vari, Richard P. 2013. Review of: Historical Biogeography of Neotropical Freshwater Fishes. Transactions of the American Fisheries Society, 142(4): 1151-1152. doi:10.1080/00028487.2013.810016

Jerep, Fernando C. and Vari, Richard P. 2013. New Species of Hemigrammocharax (Characiformes: Distichodontidae) from the Northwestern Congo River Basin. Copeia, 2013(1): 31-37. doi:10.1643/CI-12-059

Arcila, Dahiana, Vari, Richard P. and Menezes, Naércio A. 2013. Revision of the Neotropical Genus Acrobrycon (Ostariophysi: Characiformes: Characidae) with Description of Two New Species. Copeia, 2013(4): 604-611. doi:10.1643/CI-13-009

Ferraris, Carl J. and Vari, Richard P. 2013. New Species of Doumea (Siluriformes: Amphiliidae) from Coastal River Basins of the Republic of the Congo. Copeia, 2013(4): 577-581. doi:10.1643/CI-13-014

Vari, Richard P. and Ropke, Christhiana P. 2013. Familia Curimatidae. In: de Quiroz, Luiz J., Torrente-Vilara, Gislene, Ohara, Willian M., Silva Pires, Tiago H., Zuanon, Jansen and Costa Doria, Carolina R., Peixes do Rio Madeira, Volume 1. Sao Paulo: Santo Antonio, pp.112-139.

de Santana, Carlos David, Vari, Richard P. and Wosiacki, Wolmar B. 2013. The Untold Story of the Caudal Skeleton in the Electric Eel (Ostariophysi: Gymnotiformes: Electrophorus). PLoS ONE, 8(7): e68719 doi:10.1371/journal.pone.0068719

Ferraris, Carl J. J. and Vari, Richard P. 2012. Review of the African catfish genus Andersonia (Teleostei: Siluriformes). Zootaxa, 3210: 39-49.

Vari, Richard P. and Hadiaty, Renny Kurnia 2012. The endemic Sulawesi fish genus Lagusia (Teleostei: Terapontidae). Raffles Bulletin of Zoology, 60(1): 157-162.

Fernández, Luis and Vari, Richard P. 2012. New species of Trichomycterus (Teleostei: Siluriformes) from the Andean Cordillera of Argentina and the second record of the genus in thermal waters. Copeia, 2012(4): 631-636. doi:10.1643/CI-12-035

Vari, Richard P., Ferraris Jr., Carl J. and Skelton, Paul H. 2012. New species of Congoglanis (Siluriformes: Amphiliidae) from the southern Congo River basin. Copeia, 2012(4): 626-630. doi:10.1643/CI-12-029

Sidlauskas, Brian L. and Vari, Richard P. 2012. Diversity and distribution of anostomoid fishes (Teleostei: Characiformes) throughout the Guianas. Cybium, 36(1): 71-103.

Vari, Richard P., Sidlauskas , Brian L. and Le Bail, Pierre-Yves 2012. New species of Cyphocharax (Ostariophysi: Characiformes: Curimatidae) from Suriname and French Guiana and a discussion of curimatid diversity on the Guiana Shield. Cybium, 36(1): 63-69.

de Santana, Carlos D. and Vari, Richard P. 2012. New species of Adontosternarchus (Gymnotiformes, Apteronotidae) from the Rio Purus basin, Brazil. Copeia, 2012(3): 535-540. doi:10.1643/CI-11-135

Bogan, Sergio, Sidlauskas, Brian, Vari, Richard P. and Agnolin, Frederico 2012. Arrhinolemur scalabrinii Ameghino, 1898, of the late Miocene - a taxonomic journey from the Mammalia to the Anostomidae (Ostariophysi: Characiformes). Neotropical Ichthyology, 10(3): 555-560. doi:10.1590/S1679-62252012000300008

Ballen, Gustavo and Vari, Richard P. 2012. Review of the Andean armored catfishes of the genus Dolichancistrus Isbrücker (Siluformes: Loricariidae). Neotropical Ichthyology, 10(3): 499-518. doi:10.1590/S1679-62252012000300003

Mol, Jan H., Vari, Richard P., Covain, Raphael, Willink, Philip and Fisch-Muller, Sonia 2012. Annotated checklist of the freshwater fishes of Suriname. Cybium, 36(1): 293-319.

Guisande, Cástor, Pelayo-Villamil, Patricia, Vera, Manuel, Manjarrés-Hernández, Ana, Carvalho, Mónica R., Vari, Richard P., Jiménez, Lus F., Fernández, Carlos, Martínez, Paulino, Preto-Piraquive, Edgar and Granado-Lorencio, Carlos 2012. Ecological factors and diversification among Neotropical characiforms. International Journal of Ecology, 2012 doi:10.1155/2012/610419

Vari, Richard P., De Santana, Carlos D. and Wosiacki, Wolmar B. 2012. South American electric knifefishes of the genus Archolaemus (Ostariophysi, Gymnotiformes): undetected diversity in a clade of rheopiles. Zoological Journal of the Linnean Society, 165: 670-699. doi:10.1111/j.1096-3642.2012.00827.x

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