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Smithsonian National Museum of Natural History
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Department ofVertebrate Zoology

Division of Fishes

Carl Hansen
© Smithsonian Institution
bar Carole Baldwin
    Carole Baldwin
    Research Zoologist / VZ Department Chair

  • Phone: (202)633-1205
  • Fax: (202)633-0182
  • E-mail: baldwinc[at]

  • Mailing Address:
    Smithsonian Institution
    PO Box 37012, MRC 159
    Washington, DC 20013-7012

  • Shipping Address:
    Smithsonian Institution
    National Museum of Natural History
    10th and Constitution Ave, NW
    Washington, DC 20560-0159

Related Links

Journey to Galapagos


Ph.D. Marine Science, College of William and Mary
M.S. Marine Biology, College of Charleston
B.S. Biology, James Madison University

Research Interests

Diversity and evolution of coral-reef and deep-sea fishes; taxonomy of larval fishes and their utility in evolutionary studies; patterns of morphological divergence and speciation; marine conservation; public communication of marine science.

Recent Publications

Baldwin, Carole C., Tornabene, Luke, Robertson, D. Ross, Nonaka, Ai and Gilmore, R. Grant 2018. More new deep-reef basslets (Teleostei, Grammatidae, Lipogramma), with updates on the eco-evolutionary relationships within the genus. Zookeys, 729: 129-161. doi:10.3897/zookeys.729.21842

Hughes, Lily C., Ortí, Guillermo, Huang, Yu, Sun, Ying, Baldwin, Carole C., Thompson, Andrew W., Arcila, Dahiana, Betancur-R, Ricardo, Li, Chenhong, Becker, Leandro, Bellora, Nicolás, Zhao, Xiaomeng, Li, Xiaofeng, Wang, Min, Fang, Chao, Xie, Bing, Zhou, Zhuocheng, Huang, Hai, Chen, Songlin, Venkatesh, Byrappa and Shi, Qiong 2018. Comprehensive phylogeny of ray-finned fishes (Actinopterygii) based on transcriptomic and genomic data. Proceedings of the National Academy of Sciences of the United States of America, 115(24): 6249-6254. doi:10.1073/pnas.1719358115

Baldwin, Carole C., Tornabene, Luke and Robertson, D. Ross 2018. Below the Mesophotic. Scientific Reports, 8: 4920 doi:10.1038/s41598-018-23067-1

Tornabene, Luke and Baldwin, Carole C. 2017. A new mesophotic goby, Palatogobius incendius (Teleostei: Gobiidae), and the first record of invasive lionfish preying on undescribed biodiversity. Plos One, 12(5) doi:10.1371/journal.pone.0177179

Page, Lawrence M., Baldwin, Carole C., Espinosa-Pérez, Héctor, Findley, Lloyd T., Gilbert, Carter R., Hartel, Karsten E., Lea, Robert N., Mandrak, Nicholas E., Schmitter-Soto, Juan J. and Walker, H. J., Jr. 2017. Taxonomy of Gila in the Lower Colorado River Basin of Arizona and New Mexico. Fisheries, 42(9): 456-460. doi:10.1080/03632415.2017.1356108

Robertson, D. Ross, Angulo, Arturo, Baldwin, Carole C., Pitassy, Diane E., Driskell, Amy, Weigt, Lee A. and Navarro, Ignacio J. F. 2017. Deep-water bony fishes collected by the B/O Miguel Oliver on the shelf edge of Pacific Central America: an annotated, illustrated and DNA-barcoded checklist. Zootaxa, 4348(1): 1-125. doi:10.11646/zootaxa.4348.1.1

Hoeksema, Bert W., Bongaerts, Pim and Baldwin, Carole C. 2017. High coral cover at lower mesophotic depths: a dense Agaricia community at the leeward side of Curaçao, Dutch Caribbean. Marine Biodiversity, 47(1): 67-70. doi:10.1007/s12526-015-0431-8

Tornabene, Luke M., van Tassell, James L., Robertson, D. Ross and Baldwin, Carole C. 2016. Repeated invasions into the twilight zone: evolutionary origins of a novel assemblage of fishes from deep Caribbean reefs. Molecular ecology, 25(15): 3662-3682. doi:10.1111/mec.13704

Tornabene, Luke M., Robertson, D. Ross and Baldwin, Carole C. 2016. Varicus lacerta, a new species of goby (Teleostei, Gobiidae, Gobiosomatini, Nes subgroup) from a mesophotic reef in the southern Caribbean. ZooKeys, 596: 143-156. doi:10.3897/zookeys.596.8217

Tornabene, Luke, Van Tassell, James L., Gilmore, Richard G., Robertson, D. Ross, Young, Forrest and Baldwin, Carole C. 2016. Molecular phylogeny, analysis of character evolution, and submersible collections enable a new classification of a diverse group of gobies (Teleostei: Gobiidae: Nes subgroup), including nine new species and four new genera. Zoological Journal of the Linnean Society, 177(4): 764-812. doi:10.1111/zoj.12394

Sun, Ying, Huang, Yu, Li, Xiaofeng, Baldwin, Carole C., Zhou, Zhuocheng, Yan, Zhixiang, Crandall, Keith A., Zhang, Yong, Zhao, Xiaomeng, Wang, Min, Wong, Alex, Fang, Chao, Zhang, Xinhui, Huang, Hai, Lopez, Jose V., Kilfoyle, Kirk, Zhang, Yong, Ortí, Guillermo, Venkatesh, Byrappa and Shi, Qiong 2016. Fish-T1K (Transcriptomes of 1,000 Fishes) Project: large-scale transcriptome data for fish evolution studies. GigaScience, 5 doi:10.1186/s13742-016-0124-7

Baldwin, Carole C., Pitassy, Diane E. and Robertson, D. Ross 2016. A new deep-reef scorpionfish (Teleostei, Scorpaenidae, Scorpaenodes) from the southern Caribbean with comments on depth distributions and relationships of western Atlantic members of the genus. ZooKeys, 606: 141-158. doi:10.3897/zookeys.606.8590

Attaran-Farimani, Gilan, Estekani, Sanaz, Springer, Victor G., Crimmen, Oliver, Johnson, G. David and Baldwin, Carole C. 2016. Validation of the synonymy of the teleost blenniid fish species Salarias phantasticus Boulenger 1897 and Salarias anomalus Regan 1905 with Ecsenius pulcher (Murray 1887) based on DNA barcoding and morphology. Zootaxa, 4072(2): 171 doi:10.11646/zootaxa.4072.2.2

Baldwin, Carole C., Robertson, D. Ross, Nonaka, Ai and Tornabene, Luke M. 2016. Two new deep-reef basslets (Teleostei, Grammatidae, Lipogramma), with comments on the eco-evolutionary relationships of the genus. ZooKeys, 638: 45-82. doi:10.3897/zookeys.638.10455

Angulo, Arturo, Baldwin, Carole C. and Robertson, D. Ross 2016. A new species of Leptoderma Vaillant, 1886 (Osmeriformes: Alepocephalidae) from the Pacific coast of Central America. Zootaxa, 4066(4): 493-500. doi:10.11646/zootaxa.4066.4.10

Baldwin, Carole C. and Robertson, D. Ross 2015. A new, mesophotic Coryphopterus goby (Teleostei, Gobiidae) from the southern Caribbean, with comments on relationships and depth distributions within the genus. Zookeys, 513: 123-142. doi:10.3897/zookeys.513.9998

Rocha, Luiz A., Rocha, Claudia R., Baldwin, Carole C., Weigt, Lee A. and Mcfield, Melanie D. 2015. Invasive lionfish preying on critically endangered reef fish - Springer. Coral Reefs, 34(3): 803-806. doi:10.1007/s00338-015-1293-z

Anderson, William D., Jr., Johnson, G. David and Baldwin, Carole C. 2015. Review of the Splendid Perches, Callanthias (Percoidei: Callanthiidae). Transactions of the American Philosophical Society, 105(3): 1-126.

Vaslet, A., Phillips, D. L., France, Christine A. M., Feller, Ilka C. and Baldwin, Carole C. 2015. Trophic behaviour of juvenile reef fishes inhabiting interlinked mangrove-seagrass habitats in offshore mangrove islets. Journal of fish biology, 87(2): 256-273. doi:10.1111/jfb.12715

Baldwin, Carole C. and Johnson, G. David 2014. Connectivity across the Caribbean Sea: DNA Barcoding and Morphology Unite an Enigmatic Fish Larva from the Florida Straits with a New Species of Sea Bass from Deep Reefs off Curaçao. PLoS ONE, 9(5): 1-15. doi:10.1371/journal.pone.0097661

Conway, Kevin W., Baldwin, Carole C. and White, Macaulay D. 2014. Cryptic Diversity and Venom Glands in Western Atlantic Clingfishes of the Genus Acyrtus (Teleostei: Gobiesocidae). PLoS ONE, 9(5): 1-17. doi:10.1371/journal.pone.0097664

Baldwin, Carole C. and Robertson, D. Ross 2014. A new Liopropoma sea bass (Serranidae, Epinephelinae, Liopropomini) from deep reefs off Curaçao, southern Caribbean, with comments on depth distributions of western Atlantic liopropomins. ZooKeys, 409: 71-92. doi:10.3897/zookeys.409.7249

Rocha, L. A., Aleixo, A., Allen, G., Almeda, F., Baldwin, Carole C., Barclay, M. V. L., Bates, J. M., Bauer, A. M., Benzoni, F., Berns, C. M., Berumen, M. L., Blackburn, D. C., Blum, S., Bolanos, F., Bowie, R. C. K., Britz, R., Brown, R. M., Cadena, C. D., Carpenter, K., Ceriaco, L. M., Chakrabarty, P., Chaves, G., Choat, J. H., Clements, K. D., Collette, Bruce B., et al 2014. Specimen collection: An essential tool. Science, 344(6186): 814-815. doi:10.1126/science.344.6186.814

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